As a child, Shiki was taken in as an apprentice to the monster hunter Gurelli. Years later, after Gurelli"s death, Shiki returns to his former master"s town Akamaya, looking to join a monster hunter"s guild. Once in a guild, he meets Ailee, whom he discovers is Gurelli"s daughter. Shiki partners up with Ailee to take up Gurelli"s life long goal of hunting down the legendary monster Miogaruna.